A FLORISTIC INVENTORY OF THE VASCULAR PLANTS OF PRICE FALLS
Alison Mainers
McLoud High School
McLoud, Oklahoma 74851
- Abstract
- Introduction
- Study Area
- Methods
- Results and Discussion
- Table 1
- Conclusion
- Acknowledgments
- Bibliography
- Annotated Catalog
ABSTRACT
Price Falls is a small section of land located in Murray county in South central Oklahoma. The falls are found approximately five miles South of Davis. The land is owned, operated, and managed by the Independent Order of Oddfellow who have established a year round camping and recreational facility near the falls. Most of the area surveyed is on the banks of a shallow creek (Falls Creek) that feeds into the Washita River. Other areas include a travertine rock cliff face and a 45 meter stretch of limestone hillside. The goal of this project was to conduct a floristic survey of the vascular plants of Price Falls. Three collecting trips, September 1, 30, and October 28, yielded 81 genera and 42 families.
INTRODUCTION
The Oddfellow and Rebecca Camp is a family campground that holds a short camp program in the summer for youth. They offer camp and picnic sites, recreational vehicles, and swimming in the cool clean pools of Arbuckle spring water. Paying a small fee for swimming seems worth while when surrounded by many varieties of native and introduced ferns, trees, grasses and other plants. Despite its vast diversity and beauty, this area remains to be botanically unknown. No known floristic research has been conducted on the area says Chuck Conner, Vice Chairman and co-owner/care-taker of the camp with his wife, Jerriee. His knowledge of the area is also shared by the head of the botany department at Oklahoma State University, Dr. Mike Palmer. He had no knowledge on any floristic research done in the Price Falls area. My objective of this paper was to conduct a floristic survey of the vascular plants of the Price Falls area.
STUDY AREA
The Arbuckle Mountain range is located in the Grand Prarie and Cross Timbers major land resource area. The range is characterized by many natural springs, shallow streams, large river valleys, rolling limestone hills, travertine rock formations, and occasional perpendicular limestone cliffs. The Price Falls sampling area (0.44 acre) is composed of or neighbors all of these traits. The dominate soil type on the fall area is Chigley-Travertine; a well drained loamy soil that as formed in clayey material. It originated from weathered Conglomerate rock, Colluvium rock, shale, or siltstone. They are moderately alkaline to alkaline. The rock complex is dominantly limestone with slopes ranging from 0 to 90 degrees (Watterson et al., 84).
The average temperature for Murray county in the winter is 42 degrees Fahrenheit with an average daily maximum low of 30 degrees Fahrenheit. For the summer, the average temperature is 85 degrees Fahrenheit with a average maximum daily high of 95 degrees Fahrenheit. Total annual precipitation is 38 inches. Growing season is when 60 percent of this precipitation falls, April through September (Watterson et al., 84).
METHODS
Three trips were made to the fall area in monthly intervals. These dates were September 1, 30, and October 28 all in the year of 1995. The objective for each trip was to collect species from each of the seven sub-divisions the falls and surrounding areas were divided into. Identical species found in separate areas were collected to determine species abundance and apparent habitat. Both native and introduced taxa were collected. Several plants were not collected due to health reasons (Poison Ivy, Prickly Pear). Specimens were then pressed on site or field notes were made about present appearance and pressed later. The fragility of the areas surveyed was taken into consideration and no over collecting was conducted.
Identifications were made from Keys to the Flora of Oklahoma by U.T. Waterfall (1979), Forest Trees of Oklahoma by Elbert L. Little Jr. (1991), Flora of North America by the Flora of North America Editorial Committee (1993), An Annotated List of the Ferns, Fern Allies, Gymnosperms, and Flowering Plants of Oklahoma by R. John Taylor and Constance E. S. Taylor (1991), and Manual of the Vascular Plants of Texas by Donovan Correll and Marshall Johnston (1970).
RESULTS AND DISCUSSION
Floristic Summary
Based on the 284 collections made during this study, the known vascular flora of the Price Falls sampling area consists of 103 species and non-specific taxa within 81 genera and 42 families; 9 species are non-native. A statistical summary is given in Table 1 and an annotated list is included. (See addendum)
Two families, Poaceae (Gramineae) and Asteraceae (Compositae), make up almost a third of the flora with Poaceae at 18 species and Asteraceae at 14 species. Other large families include Fagaceae and Ulmaceae (6 each), and Apiaceae (4). The largest genus is Quercus (5 taxa), followed by Aster, Carya, Celtis, Panicum, Rhus, and Solidago (3 each).
Communities
There are three different plant communities at Price Falls. These communities are subdivided into smaller, more defined communities for simpler ways of collection. The communities are subjectively defined below and based on observation.
1. Creek bed.- This area is composed of: (1) small "islands" that have been disconnected with the main creek bank by many small streamlets that have been separated by the falls, and (2) the main creek bank. These areas are occasionally flooded by Falls Creek, but not by the neighboring Washita River because the area is upland from the river. This community was subdivided into two smaller areas; above falls and below falls.
2. Limestone hillside.- Four smaller sub-communities form the limestone hillside that is located across the one- way, one-lane lower road that comes out of the Falls Creek Baptist Youth Assembly. The four sub-communities are labeled as; (1) cliff- a large limestone rock face that ranges in height from two to five meters and its surroundings, (2) 1st 15- starting at the beginning of the canopy and the end of the cliff area, this is the first 15 meters of limestone hillside that slopes downward at a 45 degree angle, (3) 2nd 15- the second 15 meters down the hillside, and (4) 3rd 15- the third 15 meters down the hillside. The last 15 meters were not surveyed due to the closeness of the highway and abundance of disturbed (the area was frequently mowed down) plant life.
3. Travertine rock face.- This plant community was also called "on the falls". It was found on the travertine cliff, measuring 6 meters in height, that Falls Creek flows off of.
These communities are listed in the annotated list under the taxa that was found in them. Table 1.
Summary of the Price Falls Flora (Joyner et al.,94) Species and Lesser Taxa Families Genera Native Introduced Total Pteridophytes 4 7 7 - 7 Gymnosperms 1 1 2 - 2 Angiosperms - - - - - Momocots 3 18 17 5 22 Dicots 34 55 68 4 72 Totals 42 81 94 9 103 CONCLUSION
The objective of this study was to conduct a floristic survey of the vascular plants of Price Falls. In concluding the study, my sampling area was found to contain 42 families, 81 genera, and 103 individual species. This is an incredible diversity for such a small area (0.44 acre). The immense variety of ferns for an area less than half an acre proves how botanically rich this site is. Keeping this area and other areas like this one from commercialization and pollution depends greatly on the owner, but the responsibility is still ours to push for the preservation of these rare and magnificent places of Oklahoma.
Acknowledgments
I would like to thank my family for outstanding patience, Mrs. Courkamp and the G/T class for assistance and entertainment in long hours, and Mr. Smith for giving his all and them some.
Bibliography
- Correll, Donovan and Marshall Johnston. Manual of the Vascular Plants of Texas. Texas Research Foundation: Rener, Texas, 1970.
- Flora of North America. Oxford University Press: New York, 1993.
- Joyner, J.M. and E.W. Chester. "The Vascular Flora of Cross Creeks National Wildlife Refuge, Stewart County, Tennessee". Castanea. 59(2). 117-145.1994.
- Little, E.L. Jr. Forest Trees of Oklahoma. Oklahoma Forestry Services State Dept. of Agriculture.1991.
- Smith, Bruce. "Floristic Inventory of the McCurtain County (Oklahoma) Wilderness Area".1995.
- Taylor, R.J. and E.S. An Annotated List of the Ferns, Fern Allies, Gymnosperms, and Flowering Plants of Oklahoma. Compiled at Southeastern Oklahoma University. Durant,1991.
- Waterfall, U.T. Keys to the Flora of Oklahoma.1979.
- Watterson, A. Jr., V.A. Boggard, and G.E. Moebus. Soil Survey of Murray County, Oklahoma. Soil Conservation Service.1984.
ANNOTATED CATALOG
The catalog is divided into major groups: Pteridophyta (ferns and fern allies), Gymnosperms (non-flowering seed plants), and Angiospermae (flowering plants), which are subdivided into the Monocotylodonae and the Dicotyledonae. Within each group, families, genera, and species are arranged alphabetically. Non-native taxa are indicated by an asterisk. Common name of species is given next, followed by community type(s) where the taxon occurs. Subjective observations of abundance is given fourth based on the following scheme:
- abundant-found in community in large quantities;
- frequent-encountered in community listed, but not always in large quantities;
- occasional-found often in community, but rarely in large numbers;
- infrequent-found sparsely scattered in the stated community, usually in small numbers;
- rare-found rarely in stated community.
Collection numbers are stated last.
- Pteridophyta
- Aspleniaceae - Spleenwort Family
- Asplenium platyneuron (L.) Oakes. Ebony Spleenwort. Limestone hillside;frequent. 33,116,260,265.
- Dryopteridaceae - Wood Fern Family
- Cystopteris tenuis Michx. Mackay's brittle fern. Creek bed, travertine rock face;frequent. 83,84,99,162,225,281.
- Woodsia obtusa (Spreng) Torr. Blunt-lobed cliff fern. Limestone hillside;occasional. 116,234,239.
- Equisetaceae - Horsetail Family
- Equisetum spp. Creek bed; 71,143.
- Pteridaceae - Maidenhair Family
- Adiantum capillus-veneris L. Venus Fern. Creek bed, travertine rock face;abundant. 79,84,99,161,170,190,222, 225,278.
- Argyrochosma dealbata (Pursh) Windham, American Fern, Cloak Fern. Limestone hillside;rare. 106.
- Pellaea atropurpurea (L.) Link. Purple Cliff Break. Creek bed, limestone hillside, travertine rock face;abundant. 25, 27,60,78,99,105,106,121,194,234,239,260,265.
- Gymnospermae
- Cupressaceae - Cypress Family
- Juniperus virginiana L. Eastern Red Cedar. Limestone hillside, travertine rock face;frequent. 22,77,236,257,264, 266.
- Juniperus ashei Buckh. Ash Juniper
- Angiospermae:Monocotyledonae
- Cyperaceae - Sedge Family
- Carex spp. Creek bed;rare. 61
- Liliaceae - Lily Family
- Nothoscordum bivalve (L.) Britt. Crow poison. Limestone hillside;infrequent. 114
- Smilax bona-nox L. Greenbriar. Creek bed;occasional. 166,223
- Smilax glauca Walt. Greenbriar. Creek bed;occasional. 214
- Poaceae (Gramineae) - Grass Family
- Andropogon scoparius Michx. Little bluestem. Limestone hillside;occasional. 11.
- Eragrostis spectabilis (Pursh.) Steud. Love Grass. Creek bed, limestone hillside;frequent. 32,34,51,55,155.
- *Bromus japonicus Thunb. ex Murr. Japanese brome. Creek bed, limestone hillside;occasional. 10,180.
- Bromus pubescens Muhl. ex Willd. Canada brome. Limestone hillside;frequent. 39,46,52.
- Cenchrus incertus M.A. Curtis. Sandbur, Grassbur. Limestone hillside;occasional. 12.
- Chasmanthium latifolium (Michx.) Yates. Inland seaotes. Creek bed;abundant. 65,101,138,164,189,193,210,212.
- *Cynodon dactylon (L.) Pers. Travertine rock face;infrequent. 88.
- Elymus virginicus L. Virginia wild rye. Limestone hillside, travertine rock face;abundant. 6,21,40,112,243,255,279.
- *Leersia virginica Willd. Whitesrass. Creek bed, travertine rock face;infrequent.191,278.
- Muhlenbergia brachyphylla Bush. Bush's muhly, Nodding muhly. Limestone hillside;frequent. 119.
- Panicum spp. Creek bed;infrequent. 207.
- Panicum dichotomum L. Fall panicum. Creek bed, limestone hillside;abundant. 19,26,44,104,167,186,202,225,270,277.
- Panicum virgatum L. Fall switchgrass. Creek bed;occasional. 64,95.
- *Phalaris canariensis L. Canarygrass. Limestone hillside;infrequent. 108.
- Poa spp. Travertine rock face;rare. 87.
- Setaria geniculata (Lam.) Beauv. Knotroot bristlegrass, Foxtail. Creek bed, limestone hillside;abundant. 92,97,258.
- *Sorghum halapense (L.) Pers. Johnson grass. Creekbed;occasional. 94.
- Tridens flavus (L.) Hitchc. Purpletop. Creek bed, limestone hillside;abundant. 24,183,200,211,224,226,235,238.
- Dicotyledonae
- Acanthaceae - Acanthus Family
- Justicia americana (L.) Vahl. Denseflowered waterwillow. Creek bed;infrequent. 203.
- Anacardiaceae - Cashew Family
- Rhus aromatica Ait. Fragrant sumac. Limestone hillside; abundant. 3,5,268.
- Rhus glabra L. Smooth sumac. Limestone hillside;frequent. 1.
- Rhus copallina L. var. lanceolata Gray. Winged or Dwarfed sumac. Limestone hillside;occasional. 110.
- Apiaceae (Umbelliferae) - Parsley or Carrot Family
- Cicuta maculata L. Water hemlock, Spotted cowbane. Creek bed; occasional. 178,184.
- Eurytaenia texana T.& G. Texas spreadwing. Creek bed; occasional. 72,100,199.
- Ptilimnium nuttallii (DC.) Britt. Nuttall's Mockbishopweed. Limestone hillside;infrequent. 242.
- Sanicula canadensis L. Canada Sanicula, Snakeroot. Creek bed, limestone hillside;occasional. 7,70,240.
- Asteraceae (Compositae) - Aster, Daisy, Sunflower Family
- Ambrosia trifida L. Giant Ragweed, Horseweed. Limestone hillside;infrequent. 137.
- Aster anomalus Engelm. Aster. Creek bed;infreqent. 166,187.
- Aster azureus Lindl. Blue Aster, Azure Aster. Creek bed; infrequent. 204.
- Aster patens Ait. Spreading or Skydrop Aster. Creek bed, limestone hillside;infrequent. 217,232,259.
- Bidens vulgata Greene. Beggarstick, Sticktight. Limestone hillside;frequent. 118.
- Eupatorium coelestinum L. Mist Flower. Creek bed, travertine rock face;abundant. 76,157,165,174,196,201.
- Eupatorium purpureum L. Greenstemmed Joe-Pye Weed. Creek bed, limestone hillside;abundant. 58,103,117,147,167,197, 216,220,273.
- Helianthus hirsutus Raf. Sawtooth Sunflower. Limestone hillside;infrequent. 127,133.
- Rudbeckia hirta L. Blackeyed Susan. Creek bed; occasional. 73.
- Senecio obovatus Muhl. ex Willd. Squaweed, Roundleaf Groundsel. Creek bed;infrequent. 198.
- Solidago spp. limestone hillside, travertine rock face;occasional. 38,89.
- Solidago canadensis L. Comon Prarie Golderod. Creek bed; infrequent. 218.
- Solidago rigida L. Stiff Prarie Goldenrod. Limestone hillside;frequent. 57,126.
- *Taraxacum erythrospermum Andrz. ex Bess. Red-seed Dandelion. Limestone hillside;infrequent. 107.
- Betulaceae - Birch Family
- Ostrya virginiana (Mill.) Koch. Eastern Hop Hornbean, Ironwood. Travertine rock face;rare. 85.
- Bignoniaceae - Bignonia Family
- *Catalpa speciosa (Warder ex. Barney) Engelm. Cigar-tree, Northern Catalpa. Creek bed, travertine rock face;frequent. 80,154,168,175,185,219.
- Caprifoliaceae - Honeysuckle Family
- *Lonicera japonica Thunb. Japanese Honeysuckle. Creek bed, travertine rock face;abundant. 75,89,90,192.
- Symphoricarpos obiculatus Moench. Coralberry, Buckbrush. Creek bed, limestone hillside;abundant. 4,17,20,208,229.
- Viburnum rufidulum Raf. Rusty Blackhaw. Creek bed;frequent. 179.
- Caryophyllaceae - Pink Family
- Leoflingia texana Hook. Spreading Leoflingia. Limestone hilside;occasional. 8,241.
- Cornaceae - Dogwood Family
- Cornus drummondii Mey. Rough-leaved Dogwood. Creek bed, limestone hillside;frequent. 35,36,123,136,160.
- Ebenaceae - Ebony Family
- Diospyros virginiana L. Persimmon. Limestone hillside; infrequent. 37.
- Euphorbiaceae - Spurge Family
- Croton capitatus Michx. Woolly Croton, Hogwort. Limestone hillside;infrequent. 244.
- Euphorbia dentata Michx. Toothed Spurge. Limestone hillside; occasional. 113.
- Fabaceae (Leguminosae) - Pea, Bean Family
- Cercis canadensis L. Redbud. Creek bed, limestone hillside; frequent. 28,53,63,269.
- Desmodium nudiflorum (L.) DC. Tick Trefoil. Creek bed; infrequent. 93,102.
- Fagaceae - Beech, Oak Family
- Castanea ozarkensis Ashe. Ozark Chinkapin. Creek bed, limestone hillside;frequent. 18,56,215.
- Quercus durandii Buckl. Short-lobe Oak. Limestone hillside; frequent. 9,31,115,271.
- Quercus macrocarpa Michx. Bur Oak. Creek bed, limestone hillside;frequent. 59,144,176,177,188,209,230,231.
- Quercus muhlenbergii Engelm. Chinquapin Oak, Chestnut Oak. Creek bed, limestone hillside;frequent. 66,122,263,277.
- Quercus shumardii Buckl. Shumard's or Southern Red Oak. Limestone hillside;occasional. 30,276.
- Quercus velutina Lam. Black Oak. Limestone hillside; abundant. 54,128,131,132,267,275.
- Haloragidaceae - Water Milfoil Family
- Myriophyllum heterophyllum Michx. Coontail, Water Milfoil. Creek bed;infrequent. 178.
- Juglandaceae - Hickory, Walnut Family
- Carya illinoensis (Wang.) Koch. Pecan. Limestone hillside; occasional. 42.
- Carya texana Buckl. Black Hickory. Creek bed, limestone hillside;frequent. 120,134,151,261.
- Carya tomentosa (Poir.) Nutt. Mockernut Hickory. Limestone hillside;occasional. 49.
- Lamiaceae (Labiatae) - Mint Family
- *Perilla frutescens (L.) Britt. Common Perilla. Travertine rock face;infrequent. 169,171,172.
- Loasaceae - Loasa Family
- Mentzelia oligosperma Nutt. ex Sims. Stickleaf Mentzelia. Limestone hillside;infrequent. 13,14.
- Mimosaceae
- Albizia julibrissen Durazz. Mimosa, Baby's Breath. Creek bed, limestone hillside;infrequent. 137,159,166.
- Moraceae - Mulberry Family
- Braussonetia papyrifera (L.) Vent. Paper Mulberry. Creek bed, limestone hillside;frequent. 17,47,129,177.
- Morus alba L. White Mulberry. Creek bed;infrequent. 96.
- Morus rubra L. Red Mulberry. Creek bed;occasional. 152.
- Nyctaginaceae - Four-o' Clock Family
- Mirabilis albida (Walt.) Heimerl. White Four-o' Clock. Limestone Hillside;infrequent. 228.
- Oleaceae - Olive Family
- Fraxinus americana L. White Ash. Creek bed, limestone hillside;occasional. 15,213.
- Fraxinus pennsylvanica Marsh. Green Ash. Creek bed; infrequent. 67,139.
- Oxalidaceae - Wood Sorrel Family
- Oxalis corniculata L. Yellow Wood Sorrel, Creeping Oxalis. Limestone hillside;rare. 107.
- Plantaginaceae - Plantain Family
- Plantago spp. Creek bed;rare. 174.
- Platanaceae - Sycamore Family
- Platanus occidentalis L. Sycamore. Creek bed, travertine rock face;abundant. 62,68,74,146,150,172,173,227.
- Rhamnaceae - Buckthorn Family
- Rhamnus caroliniana Walt. Buckthorn. Creek bed, limestone hillside;frequent. 69,135,140.
- Rosaceae - Rose Family
- Crataegus reverchonii Sarg. Reverchon Hawthorn. Limestone hillside;rare. 45.
- Geum canadense Jacq. White Avens, Red Root. Limestone hillside, travertine rock face;frequent. 117,136,283.
- Rubiaceae - Madder Family
- Galium pilosum Ait. Hairy Bedstraw. Limestone hillside; infrequent. 52.
- Saliacaceae - Willow Family
- Salix nigra Marsh. Black Willow. Creek bed;infrequent. 149.
- Sapotaceae - Sapodilla Family
- Bumelia lanuginosa (Michx.) Pers. Chittamwood, Woolly Buckthorn. Creek bed, limestone hillside;frequent. 31,187, 233.
- Saxifragaceae - Saxifrage Family
- Ribes odoratum Wendl. f. Buffalo Currant, Golden Currant. Limestone hillside;occasional. 2.
- Ulmaceae - Elm Family
- Celtis laevigata Willd. Sugarberry. Creek bed;occasional. 156,168.
- Celtis occidentalis L. Hackberry. Creek bed, limestone hillside;frequent. 115,125,130,163,181.
- Celtis reticulata Torr. Neatleaf Hackberry. Limestone hillside;infrequent. 50.
- Ulmus americana L. American Elm. Creek bed, limestone hillside, travertine rock face;frequent. 85,141,153,204,262.
- Ulmus rubra Muhl. Slippery Elm. Limestone hillside; infrequent. 29.
- Verbenaceae - Vervain Family
- Verbena urticifolia L. White Vervain, Nettle Leafed Vervain. Creek bed, travertine rock face;abundant. 76,82,86,98,104, 148,182,280.
- Violaceae - Violet Family
- Viola spp. Creek bed, travertine rock face;occasional. 171, 222,280,284.
- Vitaceae - Grape Family
- Cissus incisa (Nutt.) Des Moulins. Possum Grape. Limestone hillside;occasional. 14,15,16,111.
- Parthenocissus quinquefolia (L.) Planch. Virginia Creeper. Creek bed, travertine rock face;abundant. 81,91,222.